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Today's modern filesystems are able to handle almost any desired character as part of the filename. Unfortunately this is not the case for the FAT filesystem that is commonly used on many USB thumb drives and portable music players, which is unable to use many common punctuation marks as part of the filename. GNOME 2.24 detects when you are copying files with names that contains characters that cannot be used and automatically converts these characters to a "_" without you having to rename the file yourself.